Courses

Department of Biology > Undergraduate > Courses

The courses listed below are provided by the JHU Public Course Search. This listing provides a snapshot of immediately available courses and may not be complete. Course registration information can be found on the Student Information Services (SIS) website.

Please consult the online course catalog for cross-listed courses and full course information.

Mentoring in Biology 020.441/2 Application

Mentoring in Biology provides students who have taken General Biology I and II the opportunity to mentor new students in General Biology I and II. Mentors collaborate with faculty on how to lead effective sessions, create problem sets, and generally help students understand difficult concepts and principles in biology. Mentors must have a firm command of the topics covered in biology and must meet with both faculty and students throughout the course of the semester. Students must fill out the application.

Course # (Section)	Title	Day/Times	Instructor	Location	Term	Course Details
AS.001.154 (01)	FYS: Phage Hunters - Discovering novel bacteriophages	T 12:00PM - 2:30PM	Fisher, Emily J	UG Teaching Lab (UTL) G72	Fall 2025	We often think of bacteria in the context of dangerous or annoying infections. However, bacteria themselves can be infected by even smaller and more abundant entities: viruses called bacteriophages. This First-Year Seminar will combine readings and discussion of the fundamental biology of phages and their role in controlling populations of bacteria, with lab work to discover new phages from the Johns Hopkins campus. Phages identified in this class will be added to the Science Education Alliance’s archive which is comprised of phages from over 100 academic institutions worldwide and is a resource for phage biologists and physicians directly involved in developing phages as a treatment for disease.
AS.001.186 (01)	FYS: Tuberculosis	MW 4:30PM - 5:45PM	Horner, Robert D	UG Teaching Lab (UTL) 189	Fall 2025	In the age of Molecular Biology, DNA sequencing allows the identification of genes. Biochemical assays allow the measurement of gene expression. Reverse transcriptase and PCR are used to determine the RNA made by activated genes. These tools allow the study of disease organisms on the molecular level with emphasis on particular genes, known as virulence genes, which enable the disease organisms to attack the human body. This First-Year Seminar will introduce students to the disease tuberculosis, to human innate and adaptive immune systems and to the molecular biology of Mycobacterium tuberculosis, an intracellular pathogen which infects humans and manipulates the human immune response to escape detection and elimination. We will even grow cultures of Mycobacterium smegmatum, M. tb’s harmless relative. In addition, millions worldwide have tuberculosis, and this disease is a case study in the measures that are being used to control the spread of an epidemic disease. Students will learn through by readings from books such as The White Plague by Rene’ and Jean Dubos, The Microbe Hunters by Paul de Kruif, The Aetiology of Tuberculosis by Robert Koch and Fever by John Fuller, videos, class interactions and discussions, and after the introductory portion of the course, by presentations of research papers about M. tuberculosis pathogenesis and treatment. Finally, we will explore recent genomic research that has yielded specific Mtb peptides that give evidence of being the basis of first effective vaccine for tuberculosis.
AS.020.151 (01)	General Biology I	MW 12:00PM - 1:15PM	Pearlman, Rebecca Shari; Roberson, Christov	Mudd 26	Fall 2025	This course is an introduction to biology from an evolutionary, molecular and cellular perspective. Specific topics and themes include evolutionary theory, the structure and function of biological molecules, mechanisms of harvesting energy, cell division, classical genetics and gene expression. This course is offered in fall and summer terms only.
AS.020.151 (02)	General Biology I	TTh 12:00PM - 1:15PM	Roberson, Christov	Hodson 110	Fall 2025	This course is an introduction to biology from an evolutionary, molecular and cellular perspective. Specific topics and themes include evolutionary theory, the structure and function of biological molecules, mechanisms of harvesting energy, cell division, classical genetics and gene expression. This course is offered in fall and summer terms only.
AS.020.153 (01)	General Biology Laboratory I	M 1:30PM - 4:20PM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Fall 2025	This course reinforces the topics covered in AS.020.151. Students participate in a semester-long project, identifying bacteria from Homewood campus soils using molecular biology techniques. Other laboratory exercises cover aspects of evolution, genomics and biochemistry. Cross-listed with Behavioral Biology. Student must have enrolled in AS.020.151 either this term or in past terms. Students who have credit for AP Biology but take General Biology Lab I will lose four credits of AP Biology credit. Cross-listed with Behavioral Biology. This course is offered in fall terms only.
AS.020.153 (02)	General Biology Laboratory I	T 1:30PM - 4:20PM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Fall 2025	This course reinforces the topics covered in AS.020.151. Students participate in a semester-long project, identifying bacteria from Homewood campus soils using molecular biology techniques. Other laboratory exercises cover aspects of evolution, genomics and biochemistry. Cross-listed with Behavioral Biology. Student must have enrolled in AS.020.151 either this term or in past terms. Students who have credit for AP Biology but take General Biology Lab I will lose four credits of AP Biology credit. Cross-listed with Behavioral Biology. This course is offered in fall terms only.
AS.020.153 (03)	General Biology Laboratory I	W 1:30PM - 4:20PM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Fall 2025	This course reinforces the topics covered in AS.020.151. Students participate in a semester-long project, identifying bacteria from Homewood campus soils using molecular biology techniques. Other laboratory exercises cover aspects of evolution, genomics and biochemistry. Cross-listed with Behavioral Biology. Student must have enrolled in AS.020.151 either this term or in past terms. Students who have credit for AP Biology but take General Biology Lab I will lose four credits of AP Biology credit. Cross-listed with Behavioral Biology. This course is offered in fall terms only.
AS.020.153 (04)	General Biology Laboratory I	Th 1:30PM - 4:20PM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Fall 2025	This course reinforces the topics covered in AS.020.151. Students participate in a semester-long project, identifying bacteria from Homewood campus soils using molecular biology techniques. Other laboratory exercises cover aspects of evolution, genomics and biochemistry. Cross-listed with Behavioral Biology. Student must have enrolled in AS.020.151 either this term or in past terms. Students who have credit for AP Biology but take General Biology Lab I will lose four credits of AP Biology credit. Cross-listed with Behavioral Biology. This course is offered in fall terms only.
AS.020.153 (05)	General Biology Laboratory I	F 1:30PM - 4:20PM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Fall 2025	This course reinforces the topics covered in AS.020.151. Students participate in a semester-long project, identifying bacteria from Homewood campus soils using molecular biology techniques. Other laboratory exercises cover aspects of evolution, genomics and biochemistry. Cross-listed with Behavioral Biology. Student must have enrolled in AS.020.151 either this term or in past terms. Students who have credit for AP Biology but take General Biology Lab I will lose four credits of AP Biology credit. Cross-listed with Behavioral Biology. This course is offered in fall terms only.
AS.020.153 (06)	General Biology Laboratory I	T 9:00AM - 11:50AM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Fall 2025	This course reinforces the topics covered in AS.020.151. Students participate in a semester-long project, identifying bacteria from Homewood campus soils using molecular biology techniques. Other laboratory exercises cover aspects of evolution, genomics and biochemistry. Cross-listed with Behavioral Biology. Student must have enrolled in AS.020.151 either this term or in past terms. Students who have credit for AP Biology but take General Biology Lab I will lose four credits of AP Biology credit. Cross-listed with Behavioral Biology. This course is offered in fall terms only.
AS.020.161 (01)	Current Events in Biology I	T 12:00PM - 12:50PM	Pearlman, Rebecca Shari	Gilman 55	Fall 2025	In this lively and collaborative course, students discuss current events and controversies in biology ranging from bioterrorism to the health of the Chesapeake Bay. This course is offered in fall terms only.
AS.020.304 (01)	Molecular Biology	TTh 10:30AM - 11:45AM	Broderick, Nichole; De Luca, Chiara; Fisher, Emily J	Hodson 110	Fall 2025	This course will focus on the ways that nucleic acids direct the synthesis of nucleic acids and proteins. Emphasis will be on modern techniques to study these fundamental processes and important biological molecules. This course fulfills a core requirement for biology majors and molecular and cellular biology majors. This course does not fulfill the elective requirement for biology or molecular and cellular biology majors. This course is offered in fall terms only.
AS.020.305 (01)	Biochemistry	MW 12:00PM - 1:15PM	Hilser, Vincent; Tifft Oshinnaiye, Katie Elizabeth	Hodson 110	Fall 2025	The molecules responsible for the life processes of animals, plants, and microbes will be examined. The structures, biosynthesis, degradation, and interconversion of the major cellular constituents including carbohydrates, lipids, proteins, and nucleic acids will illustrate the similarity of the biomolecules and metabolic processes involved in diverse forms of life. Sophomores, Juniors, and Seniors Only. This course is offered in fall, spring and summer terms only.
AS.020.305 (31)	JHUBiochemistry		Schildbach, Joel F	Online	Fall 2025	The molecules responsible for the life processes of animals, plants, and microbes will be examined. The structures, biosynthesis, degradation, and interconversion of the major cellular constituents including carbohydrates, lipids, proteins, and nucleic acids will illustrate the similarity of the biomolecules and metabolic processes involved in diverse forms of life. Sophomores, Juniors, and Seniors Only. This course is offered in fall, spring and summer terms only.
AS.020.314 (01)	The Biology of Disease	MW 3:00PM - 4:15PM	Tifft Oshinnaiye, Katie Elizabeth	Mudd 26	Fall 2025	Explore the current understanding of the biology of diseases in this upper-level elective! Each week, a new faculty member will present one class in a lecture style, followed by one class in an interactive discussion style. The faculty member will describe a disease and the fundamental biology relating to that disease and discuss the current state of the field, how their research influenced understanding of the disease, and progress towards treatments. The topics will build upon the basic concepts covered in genetics, cell biology, and molecular biology, and introduce topics related to biochemistry and developmental biology. The class will discuss a wide range of diseases including vision disorders, neurodegenerative diseases, and cancer. Class assessment will be based on homework involving asking questions about the seminar, writing brief summaries of seminars and discussions, and a final project related to topics and techniques from the semester. Open to juniors and seniors. This course is offered in fall terms only.
AS.020.315 (01)	Biochemistry Project lab	M 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 172	Fall 2025	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (02)	Biochemistry Project lab	M 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 174	Fall 2025	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (03)	Biochemistry Project lab	T 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 172	Fall 2025	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (04)	Biochemistry Project lab	T 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 174	Fall 2025	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (05)	Biochemistry Project lab	W 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 172	Fall 2025	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (06)	Biochemistry Project lab	W 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 174	Fall 2025	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (07)	Biochemistry Project lab	Th 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 172	Fall 2025	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (08)	Biochemistry Project lab	Th 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 174	Fall 2025	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (09)	Biochemistry Project lab	F 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 172	Fall 2025	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (10)	Biochemistry Project lab	F 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 174	Fall 2025	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.316 (01)	Cell Biology Lab	M 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 184	Fall 2025	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (02)	Cell Biology Lab	T 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 184	Fall 2025	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (03)	Cell Biology Lab	W 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 184	Fall 2025	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (04)	Cell Biology Lab	Th 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 184	Fall 2025	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (05)	Cell Biology Lab	F 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 184	Fall 2025	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.317 (01)	Great Experiments in Biology	T 2:30PM - 5:00PM	Johnston, Robert J; Kim, John	UG Teaching Lab (UTL) G89	Fall 2025	In this course, we discuss classic experiments in the history of biology and compare them to modern studies. Each week, students will present two papers, a classic paper selected by the instructors and a modern paper selected by the students. Grading will be based on student presentations, short student summaries of papers, and class participation.
AS.020.325 (01)	Scientific Research Process and Practice	MW 3:00PM - 4:15PM	Fisher, Emily J	Croft Hall G02	Fall 2025	This course is an introduction to doing great research and is designed to prepare students for successful independent research in biology. Students will learn how to find, read, and understand scientific papers, develop scientific questions, and effectively communicate their knowledge and the significance of their research in the format of a written proposal. Students will complete a short project on protein structure/interactions using computational methods to provide preliminary data for the written proposal. This course is taught by multiple co-instructors.
AS.020.331 (01)	Human Genetics	TTh 10:30AM - 11:45AM	Hedgecock, Edward M	Mudd 26	Fall 2025	Will examine the growing impact of human genetics on the biological sciences, on law and medicine, and on our understanding of human origins. Topics include structure and evolution of human genome, genetic and physical mapping of human chromosomes, molecular genetics of inherited diseases and forensic genetics. This course is offered in fall terms only
AS.020.340 (01)	Developmental Genetics Lab	T 1:30PM - 5:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 284	Fall 2025	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (02)	Developmental Genetics Lab	W 1:30PM - 5:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 284	Fall 2025	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (03)	Developmental Genetics Lab	W 1:30PM - 5:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 282	Fall 2025	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (04)	Developmental Genetics Lab	W 5:30PM - 9:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 284	Fall 2025	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (05)	Developmental Genetics Lab	W 5:30PM - 9:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 282	Fall 2025	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (06)	Developmental Genetics Lab	Th 1:30PM - 5:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 284	Fall 2025	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (07)	Developmental Genetics Lab	Th 1:30PM - 5:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 282	Fall 2025	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.350 (01)	Introduction to Clinical Medicine	T 6:30PM - 8:50PM	Meyers, David L	Hodson 313	Fall 2025	Perm. Req’d. Post-Bac Students Only. This course is offered in fall terms only.
AS.020.351 (01)	Cancer Biology	TTh 1:30PM - 2:45PM	Hoyt, Andrew Andrew	Hodson 316	Fall 2025	While the “war on cancer” has produced modest victories with respect to clinical outcomes, our knowledge of the cellular mechanisms of cancer is now vast and represents one of the most significant scientific achievements of the past 40 years. Key aspects of cancer biology will be covered with a combination of textbook and original literature readings. Topics will include cancer cell characteristics, oncogenes, tumor suppressor genes, apoptosis, metastasis and immuno-surveillance of cancer cells. Application of our knowledge to the rational treatment of cancer will also be discussed. This course is offered in fall terms only.
AS.020.374 (01)	Comparative Physiology	TTh 12:00PM - 1:15PM	Coppola, Anna	Mergenthaler 111	Fall 2025	This class examines animal physiology from an evolutionary and comparative viewpoint. The goal is to examine the commonalities, as well as unique differences, in how various animal organisms address the necessary life functions. Topics will include metabolism, neural systems, respiration, muscle systems, water and salt homeostasis, thermal regulation, and reproduction. This course is offered in fall and spring terms only.
AS.020.383 (01)	Genetic Analysis	MWF 9:00AM - 9:50AM	Andersen, Erik	Ames 234	Fall 2025	With the advent of whole-genome sequencing, many students wonder whether genetics is “solved” because we can “find the gene” for any human trait. In this course, we will build from the first principles of genetics up to human genetics in families and populations. Our goal will be to show that genetics and genetic thinking are critical to understanding human disease and a quickly growing area of modern biology. We will cover Mendelian transmission genetics to quantitative genetics using model organisms throughout to best show proofs of principle. The course will have weekly lectures, problem-solving sessions, two midterms, and a final.
AS.020.410 (01)	Teaching and Learning in Biology	W 10:00AM - 10:50AM	Fisher, Emily J; Roberson, Christov; Tifft Oshinnaiye, Katie Elizabeth	UG Teaching Lab (UTL) 189	Fall 2025	This course is by instructor permission only and exclusively for students who are invited and accepted to be learning assistants for other Biology courses. The course will focus on discussing education and application of current best teaching practices to Biology classes.
AS.020.410 (04)	Teaching and Learning in Biology	W 10:00AM - 10:50AM	Fisher, Emily J; Tifft Oshinnaiye, Katie Elizabeth	UG Teaching Lab (UTL) 189	Fall 2025	This course is by instructor permission only and exclusively for students who are invited and accepted to be learning assistants for other Biology courses. The course will focus on discussing education and application of current best teaching practices to Biology classes.
AS.020.410 (05)	Teaching and Learning in Biology	W 10:00AM - 10:50AM	Fisher, Emily J; Tifft Oshinnaiye, Katie Elizabeth	UG Teaching Lab (UTL) 189	Fall 2025	This course is by instructor permission only and exclusively for students who are invited and accepted to be learning assistants for other Biology courses. The course will focus on discussing education and application of current best teaching practices to Biology classes.
AS.020.441 (01)	Mentoring in General Biology	M 5:00PM - 5:50PM	Pearlman, Rebecca Shari	Online	Fall 2025	To become a mentor, students must have successfully completed AS.020.151/152, must apply using the form on the Biology Dept. website ( https://bio.jhu.edu/undergraduate/courses/ ), and must be accepted by the instructors. The deadline to apply is April 15th. S/U
AS.020.503 (01)	Independent Research in Biology		Tifft Oshinnaiye, Katie Elizabeth		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (02)	Independent Research in Biology		Lee, Hey-Kyoung		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (03)	Independent Research in Biology		Horner, Robert D		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (04)	Independent Research in Biology		Van Doren, Mark		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (05)	Independent Research in Biology		Green, Rachel D		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (06)	Independent Research in Biology		Trcek, Tatjana		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (07)	Independent Research in Biology		Louder, Robert K		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (08)	Independent Research in Biology		Bhabha, Gira; Ekiert, Damian		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (09)	Independent Research in Biology		Belin, Brittany		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (10)	Independent Research in Biology		Zhao, Haiqing		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (12)	Independent Research in Biology		Gordus, Andrew George		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (14)	Independent Research in Biology		Wu, Carl		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (16)	Independent Research in Biology		Kuruvilla, Rejji		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (17)	Independent Research in Biology		Norris, Carolyn R		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (20)	Independent Research in Biology		Cunningham, Kyle W		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (21)	Independent Research in Biology		McCoy, Rajiv C		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (22)	Independent Research in Biology		Schroer, Trina A		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (23)	Independent Research in Biology		DiRuggiero, Jocelyne		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (24)	Independent Research in Biology		Hedgecock, Edward M		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (27)	Independent Research in Biology		Broderick, Nichole; Fisher, Emily J; Hoyt, Andrew Andrew; Kim, Yumi; Ludington, Will		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (28)	Independent Research in Biology		Bhabha, Gira; Fisher, Emily J; Gordus, Andrew George; McCoy, Rajiv C; Schroer, Trina A		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (29)	Independent Research in Biology		Chen, Xin		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (31)	Independent Research in Biology		Roberson, Christov		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (32)	Independent Research in Biology		Zheng, Yixian		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (33)	Independent Research in Biology		Johnston, Robert J		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (36)	Independent Research in Biology		Fisher, Emily J		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (37)	Independent Research in Biology		Kirkwood, Alfred		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (38)	Independent Research in Biology		Johnson, Eric A		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (39)	Independent Research in Biology		Kaiser, Christian M		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (40)	Independent Research in Biology		Woodson, Sarah		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (41)	Independent Research in Biology		Kim, John		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (42)	Independent Research in Biology		Greider, Carol Widney		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (43)	Independent Research in Biology		Kim, Yumi		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (45)	Independent Research in Biology		Pearlman, Rebecca Shari		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (46)	Independent Research in Biology		FAN, CHEN-MING		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (47)	Independent Research in Biology		Coppola, Anna		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (48)	Independent Research in Biology		Farber, Steven		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (49)	Independent Research in Biology		Broderick, Nichole		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (50)	Independent Research in Biology		Andersen, Erik		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (51)	Independent Research in Biology		Jimenez, Erin A		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (52)	Independent Research in Biology		He, Yuan		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (53)	Independent Research in Biology		Rangan, Kavi		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (54)	Independent Research in Biology		Hilser, Vincent		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (55)	Independent Research in Biology		Tan, Frederick		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (56)	Independent Research in Biology		Ludington, Will		Fall 2025	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.505 (01)	Internship - Biology		Tifft Oshinnaiye, Katie Elizabeth		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Consent of adviser required.
AS.020.505 (02)	Internship - Biology		Roberson, Christov		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Consent of adviser required.
AS.020.505 (03)	Internship - Biology		Zhao, Haiqing		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Consent of adviser required.
AS.020.505 (04)	Internship - Biology		Fisher, Emily J		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Consent of adviser required.
AS.020.511 (03)	Independent Study		Chen, Xin		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Perm. Req’d.
AS.020.511 (04)	Independent Study		Van Doren, Mark		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Perm. Req’d.
AS.020.511 (05)	Independent Study		Fisher, Emily J		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Perm. Req’d.
AS.020.511 (06)	Independent Study		Norris, Carolyn R		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Perm. Req’d.
AS.020.511 (08)	Independent Study		Cunningham, Kyle W		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Perm. Req’d.
AS.020.511 (11)	Independent Study		Hedgecock, Edward M		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Perm. Req’d.
AS.020.511 (12)	Independent Study		Horner, Robert D		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Perm. Req’d.
AS.020.511 (13)	Independent Study		Schildbach, Joel F		Fall 2025	An independent course of study may be pursued under the direction of an adviser on those topics not specifically listed in the form of regular courses. Perm. Req’d.
AS.020.551 (01)	Mentored Research		Horner, Robert D		Fall 2025	This courses provide BA/MS students with intensive research experience for a full academic year. Students in the program work under the direction of a research mentor on an original research project, produce a written report in the form of a thesis, and make a presentation of the work to the Biology Department. BA/MS or BS/MS candidates only.
AS.080.305 (01)	Neuroscience: Cellular and Systems I	TTh 10:30AM - 11:45AM	Fetsch, Christopher R; Kuchibhotla, Kishore V; Zhao, Haiqing	Shaffer 3	Fall 2025	(Formerly Nervous Systems I) Neuroscience: Cellular and Systems I is a fully integrated, two-semester course that surveys the cellular and molecular biology of neurons as well as the structure and function of the nervous system. Students must register for Neuroscience: Cellular and Systems II offered in the second term. Course open to JHU undergraduates only.
AS.250.351 (01)	Reproductive Physiology	T 3:00PM - 4:40PM	Zirkin, Barry R	Gilman 50	Fall 2025	Focuses on reproductive physiology and biochemical and molecular regulation of the female and male reproductive tracts. Topics include the hypothalamus and pituitary, peptide and steroid hormone action, epididymis and male accessory sex organs, female reproductive tract, menstrual cycle, ovulation and gamete transport, fertilization and fertility enhancement, sexually transmitted diseases, and male and female contraceptive methods. Introductory lectures on each topic followed by research-oriented lectures and readings from current literature.
AS.360.111 (11)	SOUL: How Microbes Move	M 1:30PM - 4:00PM	Lubin, Matt	Carnegie Institute 204	Fall 2025	Microorganisms inhabit every niche of our planet and intersect with all areas of life. From traversing dry desert sands to swimming through oceans to drilling through animal tissues, these single cells get around using a dazzling array of elaborate molecular machines. How do these tiny lifeforms pull off these feats at scales where water would feel like molasses and food sources can be millions of body-lengths away? How can brainless organisms navigate mazes, swim toward nutrients, and run from predators? We’ll read landmark papers, do simple computational modeling with virtual labs, and analyze experimental data to explore the wonderful world of microbial motility. No prior background in biology or physics is required.
AS.020.126 (01)	Microbial Mysteries: Genomics from Ground to Gut	TTh 10:30AM - 11:45AM	Tan, Frederick	UG Teaching Lab (UTL) G98	Spring 2026	Microorganisms surround us, from the microbes in our soil to the bacteria in our digestive systems. This Course-based Undergraduate Research Experience will introduce computational tools to analyze metagenomic datasets and provide students the opportunity to pursue independent projects. This course is open to all students, regardless of major. no prior computational experience necessary.
AS.020.144 (01)	Science Beyond the Headline	MW 10:00AM - 11:15AM	De Luca, Chiara	Bloomberg 276	Spring 2026	This class exposes students to an active discussion of contemporary events in biology, starting with their presentation in the media and analyzing developments and controversies. The course is aimed at stimulating interest in scientific topics and at developing critical thinking. This course is not intended for Natural Science majors.
AS.020.152 (01)	General Biology II	MW 12:00PM - 1:15PM	Pearlman, Rebecca Shari; Roberson, Christov	Mudd 26	Spring 2026	This course builds on the concepts presented and discussed in General Biology I. The primary foci of this course will be on the diversity of life and on the anatomy, physiology, and evolution of plants and animals. There will be a special emphasis on human biology.
AS.020.154 (01)	General Biology Lab II	M 1:30PM - 4:20PM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Spring 2026	This course reinforces the topics covered in AS.020.152. Laboratory exercises explore subjects ranging from evolution to anatomy and physiology. Students participate in a project using molecular biology techniques to determine whether specific foods are made from genetically engineered plants. This course is offered in spring terms only. Cross-listed with Behavioral Biology
AS.020.154 (02)	General Biology Lab II	T 1:30PM - 4:20PM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Spring 2026	This course reinforces the topics covered in AS.020.152. Laboratory exercises explore subjects ranging from evolution to anatomy and physiology. Students participate in a project using molecular biology techniques to determine whether specific foods are made from genetically engineered plants. This course is offered in spring terms only. Cross-listed with Behavioral Biology
AS.020.154 (03)	General Biology Lab II	W 1:30PM - 4:20PM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Spring 2026	This course reinforces the topics covered in AS.020.152. Laboratory exercises explore subjects ranging from evolution to anatomy and physiology. Students participate in a project using molecular biology techniques to determine whether specific foods are made from genetically engineered plants. This course is offered in spring terms only. Cross-listed with Behavioral Biology
AS.020.154 (04)	General Biology Lab II	Th 1:30PM - 4:20PM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Spring 2026	This course reinforces the topics covered in AS.020.152. Laboratory exercises explore subjects ranging from evolution to anatomy and physiology. Students participate in a project using molecular biology techniques to determine whether specific foods are made from genetically engineered plants. This course is offered in spring terms only. Cross-listed with Behavioral Biology
AS.020.154 (05)	General Biology Lab II	F 1:30PM - 4:20PM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Spring 2026	This course reinforces the topics covered in AS.020.152. Laboratory exercises explore subjects ranging from evolution to anatomy and physiology. Students participate in a project using molecular biology techniques to determine whether specific foods are made from genetically engineered plants. This course is offered in spring terms only. Cross-listed with Behavioral Biology
AS.020.154 (06)	General Biology Lab II	T 9:00AM - 11:50AM	Pearlman, Rebecca Shari	UG Teaching Lab (UTL) 272	Spring 2026	This course reinforces the topics covered in AS.020.152. Laboratory exercises explore subjects ranging from evolution to anatomy and physiology. Students participate in a project using molecular biology techniques to determine whether specific foods are made from genetically engineered plants. This course is offered in spring terms only. Cross-listed with Behavioral Biology
AS.020.162 (01)	Current Events in Biology II	T 12:00PM - 12:50PM	Pearlman, Rebecca Shari	Shriver Hall Board Room	Spring 2026	Students will discuss current events and controversies in biology, ranging from genetic engineering to nanotechnology in medicine. This course is offered in spring terms only.
AS.020.303 (01)	Genetics	TTh 12:00PM - 1:15PM	Fisher, Emily J	Hodson 110	Spring 2026	This class approaches genetics as both the study of inheritance and as an approach to answer scientific questions. We cover the fundamentals of inheritance over generations and the molecular basis of resultant phenotypes, DNA sequencing and genomics, molecular evolution, and modern genetic techniques used to understand and address medically-relevant challenges. This course has no strict prerequisites, but is an upper-level science course intended for students who have taken General Biology I or have a strong biology background. Exams may be held on Thursday mornings from 8-9am, so students should have Thursday 8-9am open on their schedules.
AS.020.305 (01)	Biochemistry	TTh 10:30AM - 11:45AM	Ludington, Will; Roberson, Christov	Mudd 26	Spring 2026	The molecules responsible for the life processes of animals, plants, and microbes will be examined. The structures, biosynthesis, degradation, and interconversion of the major cellular constituents including carbohydrates, lipids, proteins, and nucleic acids will illustrate the similarity of the biomolecules and metabolic processes involved in diverse forms of life. Sophomores, Juniors, and Seniors Only.
AS.020.306 (01)	Cell Biology	MW 12:00PM - 1:15PM	Kim, Yumi; Tifft Oshinnaiye, Katie Elizabeth	Hodson 110	Spring 2026	How the molecules of living systems are organized into organelles, cells, tissues, and organisms will be explored, as well as how the activities of all of these are orchestrated and regulated to produce “life”—a phenomenon greater than the sum of its parts. Considerable emphasis is placed on experimental approaches to answering these questions. Topics covered include biological membranes, cytoskeletal elements, cell locomotion, membrane and protein traffic, the nucleus, signal transduction, the cell cycle, the extracellular matrix, epithelial structure and function. Sophomores, juniors, and seniors only. Exams may be held on Thursday mornings from 8-9am, so students should have Thursday 8-9am open on their schedules. Recommended Course Background: (AS.020.151 or AS.020.305) or equivalent knowledge of biomolecules or AS.020.303. This course is offered in spring terms only.
AS.020.312 (01)	Introduction to the Human Brain	TTh 9:00AM - 10:15AM	Hedgecock, Edward M	Mudd 26	Spring 2026	This course explores the outstanding problem of biology: how knowledge is represented in the brain. Relating insights from cognitive psychology and systems neuroscience with formal theories of learning and memory, topics include (1) anatomical and functional relations of cerebral cortex, basal ganglia, limbic system, thalamus, cerebellum, and spinal cord; (2) cortical anatomy and physiology including laminar/columnar organization, intrinsic cortical circuit, hierarchies of cortical areas; (3) activity-dependent synaptic mechanisms; (4) functional brain imaging; (5) logicist and connectist theories of cognition; and (6) relation of mental representations and natural language.
AS.020.315 (01)	Biochemistry Project lab	M 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 172	Spring 2026	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (02)	Biochemistry Project lab	M 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 174	Spring 2026	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (03)	Biochemistry Project lab	T 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 172	Spring 2026	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (04)	Biochemistry Project lab	T 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 174	Spring 2026	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (05)	Biochemistry Project lab	W 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 172	Spring 2026	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (06)	Biochemistry Project lab	W 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 174	Spring 2026	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (07)	Biochemistry Project lab	Th 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 172	Spring 2026	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (08)	Biochemistry Project lab	Th 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 174	Spring 2026	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (09)	Biochemistry Project lab	F 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 172	Spring 2026	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.315 (10)	Biochemistry Project lab	F 1:30PM - 4:30PM	Johnson, Eric A	UG Teaching Lab (UTL) 174	Spring 2026	This research project laboratory investigates the flow of energy through biological systems using focused examination of key cellular energy-conversion processes. Students will be introduced to the broad field of biochemistry research through computational structural analysis, directed mutation, recombinant protein production, and enzymatic analysis. Participants will be trained in biochemical laboratory techniques and expected to contribute their findings to the scientific community using formal, academic communications.This course is offered in fall and spring terms only.
AS.020.316 (01)	Cell Biology Lab	M 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 182	Spring 2026	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (02)	Cell Biology Lab	M 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 184	Spring 2026	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (03)	Cell Biology Lab	T 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 182	Spring 2026	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (04)	Cell Biology Lab	T 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 184	Spring 2026	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (05)	Cell Biology Lab	W 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 182	Spring 2026	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (06)	Cell Biology Lab	W 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 184	Spring 2026	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (07)	Cell Biology Lab	Th 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 182	Spring 2026	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (08)	Cell Biology Lab	Th 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 184	Spring 2026	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (09)	Cell Biology Lab	F 1:30PM - 4:30PM	Horner, Robert D	UG Teaching Lab (UTL) 182	Spring 2026	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.316 (10)	Cell Biology Lab	F 1:30PM - 4:20PM	Horner, Robert D	UG Teaching Lab (UTL) 184	Spring 2026	The Cell Biology Laboratory will use projects with the nematode C. elegans and mouse 3T3 cells in culture to illustrate experimental systems which are used in cell biology. Light microscopy, fluorescence microscopy, RNA interference, fluorescence-activated cell sorting, Western blotting and the culture of nematodes and cells are techniques which will be used. Because we will be using growing organisms, there will be at least one week when students will have to visit the lab the day after their section meets to complete an experiment. This course is offered in fall and spring terms only.
AS.020.319 (01)	Human Genome Variation	TTh 10:30AM - 11:45AM	Cunningham, Kyle W	Bloomberg 168	Spring 2026	Human Genome Variate (HGV) exposes students to the power of genomics for understanding human evolutionary history, biological traits, and medical conditions. HGV incorporates basic population genetics, direct-to-consumer DNA tests, and emerging research on human populations and their ancestors. Social and ethical issues related to the use of genetic information are also discussed. This course is offered in spring terms only.
AS.020.321 (01)	Human Genome Variation Computational Lab	W 9:00AM - 9:50AM	McCoy, Rajiv C	Bloomberg 178	Spring 2026	This option combines the main course and computational lab components of HGV. HGV exposes students to the power of genomic studies for understanding human evolutionary history, biological traits, and genetic conditions. HGV incorporates basic population genetics, direct-to-consumer DNA tests, and emerging research on human populations and their ancestors. What does real human genomic data look like? How are these data analyzed in practice? Supplementing the main course, the computational lab component will explore public datasets and bioinformatic tools used to analyze human genomic data to better understand how patterns in these data can be used to test hypotheses about evolution and human phenotypes. This course is offered in spring terms only.
AS.020.329 (01)	Microbiology	TTh 10:30AM - 11:45AM	Broderick, Nichole; DiRuggiero, Jocelyne	Shaffer 307	Spring 2026	This course explores the physiology and genetics of microorganisms within an evolutionary and ecological framework. Concepts in microbiology will be supported by molecular studies of microbial evolution and microbial communities including that of the human microbiome. Recommended Course Background: AS.020.305 This course is offered in spring and some fall terms only.
AS.020.337 (01)	Stem Cells & the Biology of Aging & Disease	T 3:00PM - 4:40PM	Wang, Jiou; Zirkin, Barry R	Mudd 100	Spring 2026	This will be a team-taught lecture course that focuses on the properties of stem cells, their possible role in cancer (breast and prostate), stem cell aging, and the potential utilization of stem cells for therapy. Topics will include: mechanisms of stem cell renewal, stem cell potency, the impact of the stem cell niche, stem cells and the hematopoietic system, stem cells and the neural system, stem cells in the male and female gonads, induced pluripotent stem cells and cellular reprogramming, stem cell changes with aging, and ethical and policy issues in stem cell research and use. Most lectures will be research-oriented. Students will be expected to read and critically analyze current literature, with an emphasis on the experimental bases from which our current understandings derive. This course is offered in spring terms only.
AS.020.340 (01)	Developmental Genetics Lab	T 1:30PM - 5:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 284	Spring 2026	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (02)	Developmental Genetics Lab	W 1:30PM - 5:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 284	Spring 2026	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (03)	Developmental Genetics Lab	W 1:30PM - 5:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 282	Spring 2026	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (04)	Developmental Genetics Lab	W 5:30PM - 9:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 284	Spring 2026	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (05)	Developmental Genetics Lab	W 5:30PM - 9:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 282	Spring 2026	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (06)	Developmental Genetics Lab	Th 1:30PM - 5:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 284	Spring 2026	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.340 (07)	Developmental Genetics Lab	Th 1:30PM - 5:20PM	Norris, Carolyn R	UG Teaching Lab (UTL) 282	Spring 2026	CRISPR (clustered regularly-interspaced short palindromic repeat) is one of the greatest advances in biology in the past decade, providing researchers with the tools to precisely and affordably edit genomes and physicians a new tool to cure disease. However, the ability to edit plant and animal genomes, including human genomes, comes with significant ethical considerations. This course will utilize a hybrid classroom-laboratory approach to provide students with both a comprehensive knowledge of the CRISPR system and a deeper understanding of how gene function is studied. At the end of the semester, you will not only understand how CRISPR works, but also have a better understanding of the power of genetics to illuminate molecular mechanisms of protein function. This course is offered in fall, spring and some summer terms only
AS.020.347 (01)	AIDS	TTh 1:30PM - 2:45PM	Schroer, Trina A	Ames 234	Spring 2026	AIDS is the world's deadliest infectious disease. This fast-paced survey course provides an in-depth treatment of topics of relevance to AIDS. We cover the biology of human immunodeficiency virus (HIV, the infectious agent that causes AIDS), the effects of HIV on the immune system, the pharmacology of the anti-viral agents that are used to suppress HIV infection, and the ongoing quest for an HIV vaccine. Because HIV drugs cannot cure HIV-infected individuals and no HIV vaccine yet exists, we will also study the long-term consequences of HIV infection including opportunistic infections, comorbid conditions, and the HIV-related cancers Kaposi's Sarcoma and AIDS-Related lymphoma. The changing social implications of chronic HIV infection are discussed throughout the semester. Recommended Course Background: AS.020.304, AS.020.305 and AS.020.306. This course is offered in spring terms only.
AS.020.351 (01)	Cancer Biology	MW 3:00PM - 4:15PM	Hoyt, Andrew Andrew	Shriver Hall 104	Spring 2026	While the “war on cancer” has produced modest victories with respect to clinical outcomes, our knowledge of the cellular mechanisms of cancer is now vast and represents one of the most significant scientific achievements of the past 40 years. Key aspects of cancer biology will be covered with a combination of textbook and original literature readings. Topics will include cancer cell characteristics, oncogenes, tumor suppressor genes, apoptosis, metastasis and immuno-surveillance of cancer cells. Application of our knowledge to the rational treatment of cancer will also be discussed. This course is offered in fall terms only.
AS.020.363 (01)	Developmental Biology	MWF 10:00AM - 10:50AM	Jimenez, Erin A; Norris, Carolyn R	Mudd 26	Spring 2026	This class will explore the development of animals from a single fertilized egg into a fully formed organism. We will emphasize experimental methods to understand the molecular mechanisms controlling development. This course is offered in spring terms only.
AS.020.364 (01)	Molecular and Cellular Mechanisms of Reproduction	M 3:00PM - 4:40PM	Wright, Bill W; Zirkin, Barry R	Mudd 100	Spring 2026	This course will address current research in the cellular and molecular biology of fundamental reproductive processes. The topics covered will vary from year to year, based on current issues in the scientific literature. The focus will be on cellular and molecular mechanisms involved in the synthesis and actions of hormones, gametogenesis, fertilization, pathologies of the reproductive tracts, developmental origins of reproductive health and disease, contraception, and infertility. The emphasis will be on defining cellular and molecular mechanisms that regulate reproductive processes, identifying the hypotheses tested in scientific papers and the strengths and limitations of experimental methods used to test the hypotheses, and evaluating and integrating data described in scientific papers. Classes will consist of a mix of lectures and student oral presentations. Recommended coursework: Reproductive Physiology. This course is offered in some fall and some spring terms only.
AS.020.366 (01)	Advances in Epigenome Sciences: Shaping the Dynamic Chromatin Landscape for Transcription	M 4:00PM - 6:30PM	Wu, Carl	Bloomberg 276	Spring 2026	Seminar on current investigations in chromatin architecture, function and dynamics at single-molecule resolution, with emphasis on the biochemical, biophysical and structural mechanisms of gene expression regulation by transcription factors, nucleosome organization and dynamics, and chromatin remodeling and modifying enzymes. Lectures, student presentations and discussion of the recent literature, with development and presentation of a mock 1-yr research proposal based on discussed publications.
AS.020.374 (01)	Comparative Physiology	TTh 12:00PM - 1:15PM	Coppola, Anna	Mergenthaler 111	Spring 2026	This class examines animal physiology from an evolutionary and comparative viewpoint. The goal is to examine the commonalities, as well as unique differences, in how various animal organisms address the necessary life functions. Topics will include metabolism, neural systems, respiration, muscle systems, water and salt homeostasis, thermal regulation, and reproduction. This course is offered in fall and spring terms only.
AS.020.375 (01)	Human Anatomy & Embryology	TTh 3:00PM - 4:30PM	Bever, Gabriel S.	Remsen Hall 1	Spring 2026	Human biology is realized through the anatomical arrangement of our cells, tissues, and organs. This course explores the details of this structural framework, emphasizing how adult anatomy is influenced by both functional demands and embryological patterning. Course content approximates that of a first-year medical school course but supports advanced study in any number of fields.
AS.020.377 (01)	Comparative Physiology Lab	M 1:30PM - 4:30PM	Coppola, Anna	UG Teaching Lab (UTL) G72	Spring 2026	This course examines the physiological principles that guide animal life processes. As a complement to the Comparative Animal Physiology lecture course, this Laboratory examines fundamental physiological principles through hands-on investigations of animal physiology using zebrafish and mussel as model systems and research-grade data acquisition systems. This course is offered in spring terms only.
AS.020.377 (02)	Comparative Physiology Lab	T 1:30PM - 4:30PM	Coppola, Anna	UG Teaching Lab (UTL) G72	Spring 2026	This course examines the physiological principles that guide animal life processes. As a complement to the Comparative Animal Physiology lecture course, this Laboratory examines fundamental physiological principles through hands-on investigations of animal physiology using zebrafish and mussel as model systems and research-grade data acquisition systems. This course is offered in spring terms only.
AS.020.410 (01)	Teaching and Learning in Biology	W 3:00PM - 3:50PM	De Luca, Chiara; Fisher, Emily J; Roberson, Christov; Tifft Oshinnaiye, Katie Elizabeth	UG Teaching Lab (UTL) 189	Spring 2026	This course is by instructor permission only and exclusively for students who are invited and accepted to be learning assistants for other biology courses. The course will focus on discussing education and application of current best teaching practices to biology classes.
AS.020.410 (02)	Teaching and Learning in Biology	W 3:00PM - 3:50PM	De Luca, Chiara; Fisher, Emily J; Roberson, Christov; Tifft Oshinnaiye, Katie Elizabeth	UG Teaching Lab (UTL) 189	Spring 2026	This course is by instructor permission only and exclusively for students who are invited and accepted to be learning assistants for other biology courses. The course will focus on discussing education and application of current best teaching practices to biology classes.
AS.020.410 (03)	Teaching and Learning in Biology	W 3:00PM - 3:50PM	De Luca, Chiara; Fisher, Emily J; Roberson, Christov; Tifft Oshinnaiye, Katie Elizabeth	UG Teaching Lab (UTL) 189	Spring 2026	This course is by instructor permission only and exclusively for students who are invited and accepted to be learning assistants for other biology courses. The course will focus on discussing education and application of current best teaching practices to biology classes.
AS.020.410 (04)	Teaching and Learning in Biology	W 3:00PM - 3:50PM	De Luca, Chiara; Fisher, Emily J; Roberson, Christov; Tifft Oshinnaiye, Katie Elizabeth	UG Teaching Lab (UTL) 189	Spring 2026	This course is by instructor permission only and exclusively for students who are invited and accepted to be learning assistants for other biology courses. The course will focus on discussing education and application of current best teaching practices to biology classes.
AS.020.442 (01)	Mentoring in General Biology	M 5:00PM - 5:50PM	Pearlman, Rebecca Shari; Shingles, Richard	Online	Spring 2026	This course provides students who have taken General Biology I & II the opportunity to mentor new students in General Biology I & II. Mentors collaborate with faculty on how to lead effective sessions, create study materials for students, help student teams complete team assignments, and generally help students understand difficult concepts and principles in biology. Mentors must have a firm command of the topics covered in biology and must meet with both faculty and students through the course of the semester. To become a mentor, students must have successfully completed AS.020.151/AS.020.152, must apply using the form on the Biology Department website, and must be accepted by the instructors.
AS.020.503 (01)	Independent Research in Biology		Tifft Oshinnaiye, Katie Elizabeth		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (02)	Independent Research in Biology		Lee, Hey-Kyoung		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (03)	Independent Research in Biology		Horner, Robert D		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (04)	Independent Research in Biology		Van Doren, Mark		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (05)	Independent Research in Biology		Green, Rachel D		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (06)	Independent Research in Biology		Trcek, Tatjana		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (07)	Independent Research in Biology		Hilser, Vincent		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (08)	Independent Research in Biology		Jimenez, Erin A		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (09)	Independent Research in Biology		Belin, Brittany		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (10)	Independent Research in Biology		Zhao, Haiqing		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (11)	Independent Research in Biology		Ludington, Will		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (12)	Independent Research in Biology		Gordus, Andrew George		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (14)	Independent Research in Biology		Broderick, Nichole		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (16)	Independent Research in Biology		Kuruvilla, Rejji		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (17)	Independent Research in Biology		Norris, Carolyn R		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (18)	Independent Research in Biology		Coppola, Anna		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (19)	Independent Research in Biology		Farber, Steven		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (20)	Independent Research in Biology		Cunningham, Kyle W		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (21)	Independent Research in Biology		McCoy, Rajiv C		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (22)	Independent Research in Biology		Moudrianakis, E N		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (23)	Independent Research in Biology		DiRuggiero, Jocelyne		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (24)	Independent Research in Biology		Hedgecock, Edward M		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (25)	Independent Research in Biology		Andersen, Erik		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (26)	Independent Research in Biology		Bhabha, Gira; Ekiert, Damian		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (27)	Independent Research in Biology		Broderick, Nichole; Fisher, Emily J; Hoyt, Andrew Andrew; Kim, Yumi; Ludington, Will		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. This section is for members for the class of 2027 and is administered by the class of 2027 mentoring team. Applications to apply for enrollment are available in our majors' Canvas site or by contacting an instructor.
AS.020.503 (28)	Independent Research in Biology		Bhabha, Gira; Fisher, Emily J; Gordus, Andrew George; McCoy, Rajiv C; Schroer, Trina A		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. This section is for members for the class of 2028 and is administered by the class of 2028 mentoring team. Applications to apply for enrollment are available in our majors' Canvas site or by contacting an instructor.
AS.020.503 (29)	Independent Research in Biology		Chen, Xin		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (31)	Independent Research in Biology		Roberson, Christov		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (32)	Independent Research in Biology		Zheng, Yixian		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (33)	Independent Research in Biology		Johnston, Robert J		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (35)	Independent Research in Biology		Pearlman, Rebecca Shari		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (36)	Independent Research in Biology		Fisher, Emily J		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. This section is intended for students outside the Bio or MCB majors, including pre-major freshmen. Please contact Dr. Fisher regarding enrollment.
AS.020.503 (37)	Independent Research in Biology		Kirkwood, Alfred		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (38)	Independent Research in Biology		Johnson, Eric A		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (40)	Independent Research in Biology		Woodson, Sarah		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (41)	Independent Research in Biology		Kim, John		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (43)	Independent Research in Biology		Kim, Yumi		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (45)	Independent Research in Biology		Wu, Carl		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.503 (46)	Independent Research in Biology		Rangan, Kavi		Spring 2026	Planning and conducting original laboratory investigations on biological problems, collection and analysis of data, reporting of results. Permission of full-time faculty member in Biology dept.
AS.020.506 (01)	Internship - Biology		Cunningham, Kyle W		Spring 2026	For students undertaking internships that focus on biology fields or topics. Students wishing to enroll in this course must consult with the director of undergraduate studies or their major advisor before submitting an Independent Study Request in SIS.
AS.020.506 (02)	Internship - Biology		Van Doren, Mark		Spring 2026	For students undertaking internships that focus on biology fields or topics. Students wishing to enroll in this course must consult with the director of undergraduate studies or their major advisor before submitting an Independent Study Request in SIS.
AS.020.506 (03)	Internship - Biology		Norris, Carolyn R		Spring 2026	For students undertaking internships that focus on biology fields or topics. Students wishing to enroll in this course must consult with the director of undergraduate studies or their major advisor before submitting an Independent Study Request in SIS.
AS.020.506 (04)	Internship - Biology		Tifft Oshinnaiye, Katie Elizabeth		Spring 2026	For students undertaking internships that focus on biology fields or topics. Students wishing to enroll in this course must consult with the director of undergraduate studies or their major advisor before submitting an Independent Study Request in SIS.
AS.020.506 (08)	Internship - Biology		Schildbach, Joel F		Spring 2026	For students undertaking internships that focus on biology fields or topics. Students wishing to enroll in this course must consult with the director of undergraduate studies or their major advisor before submitting an Independent Study Request in SIS.
AS.020.506 (09)	Internship - Biology		Fisher, Emily J		Spring 2026	For students undertaking internships that focus on biology fields or topics. Students wishing to enroll in this course must consult with the director of undergraduate studies or their major advisor before submitting an Independent Study Request in SIS.
AS.020.506 (10)	Internship - Biology		Roberson, Christov		Spring 2026	For students undertaking internships that focus on biology fields or topics. Students wishing to enroll in this course must consult with the director of undergraduate studies or their major advisor before submitting an Independent Study Request in SIS.
AS.020.553 (01)	Mentored Research		Horner, Robert D		Spring 2026	BA/MS candidates only.
AS.020.553 (03)	Mentored Research		Beemon, Karen L		Spring 2026	BA/MS candidates only.
AS.020.553 (04)	Mentored Research		Zhao, Haiqing		Spring 2026	BA/MS candidates only.

Courses

Mentoring in Biology 020.441/2 Application

FYS: Phage Hunters - Discovering novel bacteriophagesAS.001.154 (01)

FYS: TuberculosisAS.001.186 (01)

General Biology IAS.020.151 (01)

General Biology IAS.020.151 (02)

General Biology Laboratory IAS.020.153 (01)

General Biology Laboratory IAS.020.153 (02)

General Biology Laboratory IAS.020.153 (03)

General Biology Laboratory IAS.020.153 (04)

General Biology Laboratory IAS.020.153 (05)

General Biology Laboratory IAS.020.153 (06)

Current Events in Biology IAS.020.161 (01)

Molecular BiologyAS.020.304 (01)

BiochemistryAS.020.305 (01)

JHUBiochemistryAS.020.305 (31)

The Biology of DiseaseAS.020.314 (01)

Biochemistry Project labAS.020.315 (01)

Biochemistry Project labAS.020.315 (02)

Biochemistry Project labAS.020.315 (03)

Biochemistry Project labAS.020.315 (04)

Biochemistry Project labAS.020.315 (05)

Biochemistry Project labAS.020.315 (06)

Biochemistry Project labAS.020.315 (07)

Biochemistry Project labAS.020.315 (08)

Biochemistry Project labAS.020.315 (09)

Biochemistry Project labAS.020.315 (10)

Cell Biology LabAS.020.316 (01)

Cell Biology LabAS.020.316 (02)

Cell Biology LabAS.020.316 (03)

Cell Biology LabAS.020.316 (04)

Cell Biology LabAS.020.316 (05)

Great Experiments in BiologyAS.020.317 (01)

Scientific Research Process and PracticeAS.020.325 (01)

Human GeneticsAS.020.331 (01)

Developmental Genetics LabAS.020.340 (01)

Developmental Genetics LabAS.020.340 (02)

Developmental Genetics LabAS.020.340 (03)

Developmental Genetics LabAS.020.340 (04)

Developmental Genetics LabAS.020.340 (05)

Developmental Genetics LabAS.020.340 (06)

Developmental Genetics LabAS.020.340 (07)

Introduction to Clinical MedicineAS.020.350 (01)

Cancer BiologyAS.020.351 (01)

Comparative PhysiologyAS.020.374 (01)

Genetic AnalysisAS.020.383 (01)

Teaching and Learning in BiologyAS.020.410 (01)

Teaching and Learning in BiologyAS.020.410 (04)

Teaching and Learning in BiologyAS.020.410 (05)

Mentoring in General BiologyAS.020.441 (01)

Independent Research in BiologyAS.020.503 (01)

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