People

Below are our Professor Emeriti, Tenured And Tenure Track Faculty, Joint Appointments, Adjunct Appointments, Research Faculty, and Lecturers grouped by designation. To see the Department’s full faculty directory listed alphabetically, please visit our Biology Faculty (Alphabetical) page.

Please note: adjunct Carnegie faculty will be leaving Johns Hopkins University at the end of 2025 and are no longer accepting JHU students into their labs.

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  • Tenured and Tenure Track Faculty

  • Erik Andersen

    Erik Andersen

    Professor

    • erik.andersen@jhu.edu
    • UTL 383
    • 410-516-1282
    • Research Interests: Genetics and genomics of complex traits; host-pathogen genomics and drug resistance; evolutionary genetics
  • Nichole Broderick

    Nichole Broderick

    Assistant Professor

    • nbroder1@jhu.edu
    • Levi 236
    • Group/Lab Website
    • Research Interests: Molecular and cellular mechanisms by which the microbiome affects animal development, physiology and immunity
    • Education: PhD, University of Wisconsin-Madison
  • Xin Chen

    Xin Chen

    Professor, HHMI Investigator

    • xchen32@jhu.edu
    • 137 Levi Hall
    • 410-516-4576 | Lab 410-516-6543
    • Research Interests: Epigenetic regulation of stem cell lineages
    • Education: PhD, University of Texas at Austin
  • Kyle Cunningham

    Kyle Cunningham

    Professor

    • kwc@jhu.edu
    • 204 Bio East
    • 410-516-7844 |Lab 410-516-6638
    • Research Interests: Mechanisms of calcium signaling and cell death
    • Education: PhD, University of California, Los Angeles
  • Jocelyne DiRuggiero

    Jocelyne DiRuggiero

    Associate Professor

    • jdiruggiero@jhu.edu
    • 235 Levi Hall
    • 410-516-8498 | Lab 410-516-7744
    • Group/Lab Website
    • Research Interests: Molecular microbial ecology of extremophiles; molecular mechanisms of stress response
  • Steven Farber

    Steven Farber

    Director, CMDB Graduate Program;
    Professor, Department of Biology

    • sfarber3@jhu.edu
    • 253 Levi Hall
    • 410-246-3072
    • Group/Lab Website
    • Research Interests: Real-time imaging of lipid metabolism in live zebrafish; cell and molecular biology of lipids; identification of genes which regulate digestive organ lipid metabolism using biochemical and genetic strategies
  • Andrew Gordus

    Andrew Gordus

    Assistant Professor

    • agordus@jhu.edu
    • 207 Biology East
    • 410-516-6509
    • Group/Lab Website
    • Research Interests: Understanding how novel and innate behaviors are encoded at the cellular and genetic level
    • Education: PhD, Harvard University
  • Rachel Green

    Rachel Green

    Bloomberg Distinguished Professor

  • Edward Hedgecock

    Edward Hedgecock

    Professor

    • ehedgec1@jhu.edu
    • 007 Biology East
    • Lab 410-516-8381
    • Research Interests: Developmental genetics of the nervous system of Caenorhabditis elegans
    • Education: PhD, University of California, Santa Cruz
  • Vincent Hilser

    Vincent Hilser

    Professor and Chair

    • hilser@jhu.edu
    • Mudd 123
    • 410-516-4693 | Lab 410-516-6757
    • Group/Lab Website
    • Research Interests: protein conformational fluctuations and their role in function, adaptation and disease
    • Education: PhD, Johns Hopkins University
  • M. Andrew Hoyt

    M. Andrew Hoyt

    Professor

    • hoyt@jhu.edu
    • Mudd 206
    • 410-516-7299 |Lab 410-516-7293
    • Research Interests: Yeast chromosome segregation with emphasis on mitotic motor proteins; cell cycle regulation
    • Education: PhD, University of California, Berkeley
  • Erin Jimenez

    Erin Jimenez

    Assistant Professor

    • ejimene4@jh.edu
    • 253 Levi Hall
    • Research Interests: Gene regulatory networks in vertebrate inner ear regeneration
  • Robert Johnston

    Robert Johnston

    Associate Professor

    • robertjohnston@jhu.edu
    • 309 Mudd Hall
    • 410-516-4954
    • Research Interests: Development and gene regulation in human retinal organoids and fly eyes
    • Education: PhD, Columbia University
  • John Kim

    John Kim

    Associate Professor

    • jnkim@jhu.edu
    • 389 UTL
    • 410-516-3286 | Lab 410-516-3463
    • Group/Lab Website
    • Research Interests: Deciphering the biological function of microRNAs and other small RNAs expressed in the genome
  • Yumi Kim

    Yumi Kim

    Assistant Professor

  • Rejji Kuruvilla

    Rejji Kuruvilla

    Professor

    • rkuruvilla@jhu.edu
    • 227 Mudd Hall
    • 410-516-2366 | Lab 410-516-7641
    • Group/Lab Website
    • Research Interests: Control of neuronal development by target-derived neurotrophins
    • Education: PhD, University of Houston
  • Rajiv McCoy

    Rajiv McCoy

    Assistant Professor

    • rajiv.mccoy@jhu.edu
    • Levi 249
    • 410-516-0942
    • Group/Lab Website
    • Research Interests: Computational and statistical genomic approaches for studying human evolution and reproduction
    • Education: PhD, Stanford University
  • Evangelos Moudrianakis

    Evangelos Moudrianakis

    Professor

    • vanm@jhu.edu
    • 20 B Bio East
    • 410-516-7305 | Lab 410-516-7303
    • Research Interests: Assembly and dynamics of nucleoproteins and chromosomes; bacterial and chloroplast bioenergetics
    • Education: PhD, Johns Hopkins University
  • Ulrich Mueller

    Ulrich Mueller

    Bloomberg Distinguished Professor

  • Joel Schildbach

    Joel Schildbach

    PhD, Harvard University

    • joel@jhu.edu
    • 410-516-5966
    • Research Interests: Structural biology of bacterial conjugation
  • Michael Schatz

    Michael Schatz

    Bloomberg Distinguished Professor of Computer Science and Biology

    • mschatz@jhu.edu
    • (410) 516-5765
    • Group/Lab Website
    • Research Interests: My research interests are Genome sequencing, Comparative Genomics, Single Cell and Single Molecule Biology, Bioinformatics
  • Trina Schroer

    Trina Schroer

    Professor

    • schroer@jhu.edu
    • 203 Biology East
    • 410-516-5373 | Lab 410-516-5374
    • Research Interests: Microtubule-based motor proteins and their roles in cellular architecture
    • Education: PhD, University of California, San Francisco
  • Tatjana Trcek

    Tatjana Trcek

    Assistant Professor

    • ttrcekp1@jhu.edu
    • Mudd 301A
    • 410-516-2332 (office) 410-516-1152 (lab)
    • Group/Lab Website
    • Research Interests: Spatial organization of cellular mRNAs, RNA granules, imaging
    • Education: PhD, Albert Einstein College of Medicine
  • Mark Van Doren

    Mark Van Doren

    Professor

    • vandoren@jhu.edu
    • BioEast 16A
    • 410-516-4717 | Lab 410-516-4830
    • Group/Lab Website
    • Research Interests: Germ cell development and gonad formation
    • Education: PhD, University of California, San Diego
  • Carl Wu

    Carl Wu

    Bloomberg Distinguished Professor

    • wuc@jhu.edu
    • 387 UTL
    • 410-516-7496
    • Research Interests: Eukaryotic chromatin structure and function. Biochemistry of nucleosome remodeling and gene transcription, and live cell-single molecule imaging of histone and nuclear protein dynamics
  • Haiqing Zhao

    Haiqing Zhao

    Professor

    • hzhao@jhu.edu
    • 226 Mudd Hall
    • 410-516-7391 | Lab 410-516-7641
    • Research Interests: Function and development of olfactory sensory neurons
    • Education: PhD, Yale University

  • Joint Appointments

  • Douglas Barrick

    Associate Professor, Department of Biophysics

    • barrick@jhu.edu
    • 410-516-0409
    • Research Interests: Determination of an atomic-level description of structure and function in proteins

  • Alexis Battle

    Associate Professor, Department of Biomedical Engineering

  • Gregory Bowman

    Associate Professor, Department of Biophysics

  • Brian Camley

    Assistant Professor, Department of Physics and Astronomy

    • bcamley@jhu.edu
    • Bloomberg 243
    • 410-516-7683
    • Group/Lab Website
    • Research Interests: Cell motility, chemotaxis, collective motion, soft matter, and fluid mechanics in cell biology

  • Karen Fleming

    Professor, Department of Biophysics

    • karen.fleming@jhu.edu
    • 410-554-1244
    • Research Interests: Structure, energetics and folding of membrane proteins

  • Stephen Fried

    Assistant Professor, Department of Chemistry

    • sdfried@jhu.edu
    • Remsen 121
    • 410-516-7835
    • Research Interests: Folding and assembly of proteins and molecular machines in vivo; crosslinking mass spectrometry; ancient proteins and the origins of life; synthetic biology and directed evolution.

  • Bertrand E. Garcia-Moreno

    Professor, Department of Biophysics; KSAS Vice Dean for Natural Sciences

    • bertrand@jhu.edu
    • 410-516-4497
    • Research Interests: Biophysical chemistry of regulatory interactions in proteins, nucleic acids, and their assemblies

  • Marc M. Greenberg

    Professor, Department of Chemistry

    • mgreenberg@jhu.edu
    • New Chemistry Building 313
    • 410-516-8095
    • Research Interests: Nucleic acid damage and repair

  • Margaret Johnson

    Assistant Professor, Department of Biophysics

    • mjohn218@jhu.edu
    • 410-516-2376
    • Research Interests: Theoretical and computational modeling of protein interactions at the cellular scale

  • Alfredo Kirkwood

    Professor, Department of Neuroscience (School of Medicine)

  • Juliette Lecomte

    Professor, Department of Biophysics

  • Hey-Kyoung Lee

    Associate Professor, Department of Neuroscience

    • heykyounglee@jhu.edu
    • 410-516-5712
    • Research Interests: Cellular and molecular mechanisms of synaptic plasticity underlying memory formation

  • Steven L. Salzberg

    Bloomberg Distinguished Professor

    • salzberg@jhu.edu
    • Wyman S261
    • 410-516-8246
    • Group/Lab Website
    • Research Interests: Computational biology, genomics, DNA sequence analysis, metagenomics, sequencing technology
    • Education: PhD

  • Craig A. Townsend

    Professor, Department of Chemistry

  • Sarah Woodson

    Professor, Department of Biophysics

  • Adjunct Appointments

  • Brittany Belin

    Adjunct Assistant Professor, Department of Biology; Staff, Carnegie Institution

    • belin@carnegiescience.edu
    • On Leave: Carnegie faculty will be leaving Johns Hopkins University at the end of 2025 and are no longer accepting JHU students into their labs.
    • Group/Lab Website
    • Research Interests: Cell biology of symbiotic bacteria and their associations with plants.

  • Alex Bortvin

    Staff member, Department of Embryology, Carnegie Institution of Washington
    Adjunct Assistant Professor, Biology

    • bortvin@carnegiescience.edu
    • On Leave: Carnegie faculty will be leaving Johns Hopkins University at the end of 2025 and are no longer accepting JHU students into their labs.
    • 410-246-3034
    • Research Interests: Genetic and epigenetic controls of germ cell development and function in vertebrates

  • Phillip Cleves

    Adjunct Assistant Professor, Department of Biology; Staff, Carnegie Institution of Washington

    • cleves@carnegiescience.edu
    • On Leave: Carnegie faculty will be leaving Johns Hopkins University at the end of 2025 and are no longer accepting JHU students into their labs.
    • Research Interests: Genetic and cellular bases of coral-algal symbiosis and its breakdown due to stress

  • Chen-Ming Fan

    Staff Member, Department of Embryology, Carnegie Institution of Washington
    Adjunct Professor, Biology

    • fan@carnegiescience.edu
    • On Leave: Carnegie faculty will be leaving Johns Hopkins University at the end of 2025 and are no longer accepting JHU students into their labs.
    • 410-246-3022
    • Research Interests: Molecular patterning and embryonic induction during mouse early development

  • Joseph Gall

    Staff Member, Department of Embryology, Carnegie Institution of Washington
    Adjunct Professor, Biology

    • gall@carnegiescience.edu
    • On Leave: Carnegie faculty will be leaving Johns Hopkins University at the end of 2025 and are no longer accepting JHU students into their labs.
    • 410-246-3018
    • Research Interests: Chromosome structure and function; nucleic acids in development

  • Marnie Halpern

    Chair and Professor of Molecular and Systems Biology, the Geisel School of Medicine at Dartmouth
    Adjunct Professor, Biology

  • Christian Kaiser

    Christian Kaiser

    Associate Professor

    • kaiser@jhu.edu
    • 205 Biology East
    • 410-516-4486
    • Group/Lab Website
    • Research Interests: Single-molecule biochemistry studies of the machines and processes in protein translation, translocation, and folding
    • Education: Dr. rer. nat. (PhD), Ludwig-Maximilians-Universität, Munich; Germany Max Planck Institute of Biochemistry, Martinsried, Germany

  • William Ludington

    Staff Member, Department of Embryology, Carnegie Institution of Washington
    Adjunct Assistant Professor, Biology

    • ludington@carnegiescience.edu
    • On Leave: Carnegie faculty will be leaving Johns Hopkins University at the end of 2025 and are no longer accepting JHU students into their labs.
    • (410) 246-3003
    • Research Interests: Mechanisms of complexity in microbiome-host interactions

  • George Scangos

    CEO & Director, Vir
    Adjunct Professor, Biology (Biogen Idec.)

  • Allan Spradling

    Director, Department of Embryology, Carnegie Institution of Washington
    Adjunct Professor, Biology

    • spradling@carnegiescience.edu
    • On Leave: Carnegie faculty will be leaving Johns Hopkins University at the end of 2025 and are no longer accepting JHU students into their labs.
    • 410-246-3021
    • Research Interests: Drosophila genomics

  • Yixian Zheng

    Staff Member, Department of Embryology, Carnegie Institution of Washington
    Adjunct Professor, Biology

    • zheng@carnegiescience.edu
    • On Leave: Carnegie faculty will be leaving Johns Hopkins University at the end of 2025 and are no longer accepting JHU students into their labs.
    • 410-246-3032
    • Research Interests: Mechanisms of cell division and genome organization in development, homeostasis, and aging

  • Research Faculty

  • Ian Dobbie

    Ian Dobbie

    Research Professor and Director of the Integrated Imaging Center

    • ian.dobbie@jhu.edu
    • 410-516-3321
    • Research Interests: Building novel super-resolution fluorescence microscopes for cutting edge bioimaging studies
    • Education: PhD, Randall Institute, King’s College London

  • Ru Chih Huang

    McElroy Honorary Research Professor

    • rhuang@jhu.edu
    • 250 Levi Hall
    • 410-516-5181 | Lab 410-516-7314
    • Research Interests: Gene regulation, molecular virology, retrotransposition, and chromosomal structure and function
  • Yuan Chuan Lee

    Yuan Chuan Lee

    Academy Professor and Research Professor

    • yclee@jhu.edu
    • Levi 141 A
    • 410-516-7322
    • Research Interests: Glycobiology, glycotechnology, and glyconanotechnology

  • Lecturers

  • Anna Coppola

    Lecturer

    • anna.coppola@jhu.edu
    • Macaulay 107
    • Research Interests: Effect of dietary nutrients on metabolic regulatory mechanisms, with particular emphasis on the role of proteins and amino acids in the development of behavioral and metabolic diseases.
    • Education: Ph.D. in Pharmacology (2005), University of Naples ‘Federico II’, Italy. B.Sc. Pharmaceutical Biotechnology (2001), University of Naples ‘Federico II’, Italy.
  • Emily Fisher

    Emily Fisher

    Director of Undergraduate Studies, Associate Teaching Professor

    • efishe21@jhu.edu
    • 181 UTL
    • 410-516-8044
    • Research Interests: How and with whom students work to learn biology. Development of effective teaching practices in science communication.
    • Education: PhD. in Biology, University of North Carolina, Chapel Hill (2008); B.S. in Biochemistry and Molecular Biology, University of California Santa Cruz (2000)
  • Robert Horner

    Robert Horner

    Senior Lecturer

    • rdhorner@jhu.edu
    • 179 UTL
    • 410-516-8067 | Lab 410-516-7281
    • Education: PhD, The Johns Hopkins University

  • Eric Johnson

    Senior Lecturer

    • ejohn@jhu.edu
    • Office - Macaulay 107, Lab - UTL 172
    • 410-516-4147
    • Education: Ph.D., Johns Hopkins University
  • Carolyn Norris

    Carolyn Norris

    Senior Lecturer

    • crn@jhu.edu
    • 279 UTL
    • 410-516-8288 | Lab 410-516-4666
    • Education: PhD, University of Wisconsin
  • Rebecca Pearlman

    Rebecca Pearlman

    Senior Lecturer

    • pearlman@jhu.edu
    • 281 UTL
    • (410) 516-5479
    • Research Interests: Innovations in Undergraduate Biology Education
    • Education: PhD., University of Wisconsin; B.S. Hons., University of Michigan

  • Christov Roberson

    Lecturer

  • Richard Shingles

    Richard Shingles

    Lecturer

    • shingles@jhu.edu
    • 281 UTL
    • 410-516-4679
    • Research Interests: Membrane biochemistry, Membrane transport mechanisms, Stopped-flow fluorescence
    • Education: PhD, University of Guelph; B. Ed., University of Toronto

  • Katie Tifft

    Associate Teaching Professor; Assistant Director, Molecular and Cellular Biology Master's Program

  • Professor Emeriti

  • Karen Beemon

    Karen Beemon

    Professor Emeritus

    • klb@jhu.edu
    • 20A Biology East
    • 410-516-7289 | Lab 410-516-8236
    • Research Interests: Role of TERT and microRNAs in oncogenesis, retroviral RNA stability
    • Education: PhD, University of California, Berkeley

  • Michael Edidin

    Professor Emeritus

  • Douglas M. Fambrough

    Professor Emeritus

  • Ernesto Freire

    Ernesto Freire

    Henry Walters Professor
    Professor Emeritus

    • ef@jhu.edu
    • 114A Biology East
    • 410-516-7743 | Lab 410-516-7742
    • Research Interests: Structure-based thermodynamics of molecular recognition and function
    • Education: PhD, University of Virginia
  • Richard McCarty

    Richard McCarty

    Professor Emeritus

    • rem1@jhu.edu
    • 410-516-5356
    • Research Interests: Structure, mechanism, and regulation of the chloroplast ATP synthase; chloroplast metabolite transport
  • Robert Schleif

    Robert Schleif

    Professor Emeritus

    • schleif@jhu.edu
    • 239 Mudd Hall
    • 410-516-5206
    • Research Interests: Protein-DNA interactions and regulation of gene activity
    • Education: PhD, University of California, Berkeley

  • Allen Shearn

    Professor Emeritus

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